References
  -  A.M. Saunders, M. Albertsen, J. Vollertsen, P.H. Nielsen, The
    activated sludge ecosystem contains a core community of
    abundant organisms, ISME J., 10 (2016) 111–119. 
-  A. Sengar, A. Aziz, I.H. Farooqi, F. Basheer, Development of
    denitrifying phosphate accumulating and anammox microorganisms
    in anaerobic hybrid reactor for removal of nutrients
    from low strength domestic sewage, Bioresour. Technol.,
    83 (2018) 559–568. 
-  D. Li, X. Liang, Y. Jin, C. Wu, R.Q. Zhou, Isolation and
    nitrogen removal characteristics of an aerobic heterotrophic
    nitrifying-denitrifying bacterium, Klebsiella sp. TN-10, Biochem.
    Biotechnol., 188 (2019) 540–554. 
-  H. Liu, Q. Wang, Y. Sun, K. Zhou, W. Liu, Q. Lu, C. Ming,
    X. Feng, J. Du, X. Jia, J. Li, Isolation of a non-fermentative
    bacterium, Pseudomonas aeruginosa, using intracellular carbon
    for denitrification and phosphorus accumulation and relevant
    metabolic mechanisms, Bioresour. Technol., 211 (2016) 6–15. 
-  Q. Jia, H. Xiong, H. Wang, H. Shi, X. Sheng, R. Sun, G. Chen,
    Production of polyhydroxyalkanoates (PHA) by bacterial
    consortium from excess sludge fermentation liquid at
    laboratory and pilot scales, Bioresour. Technol., 171 (2017)
    159–167. 
-  M. Carvalheira, A. Oehmen, G. Carvalho, M.A.M. Reis, Survival
    strategies of polyphosphate accumulating organisms and
    glycogen accumulating organisms under conditions of low
    organic loading, Bioresour. Technol., 172 (2014) 290–296. 
-  A. Chojnacka, M.K. Błaszczyk, P. Szczęsny, K. Nowak,
    M. Sumińska, K. Tomczyk-Żak, U. Zielenkiewicz, A. Sikora,
    Comparative analysis of hydrogen-producing bacterial
    biofilms and granular sludge formed in continuous cultures
    of fermentative bacteria, Bioresour. Technol., 102 (2014)
    10057–10064. 
-  J. Fang, P.-d. Sun, S.-j. Xu, T. Luo, J.-q. Lou, J.-y. Han, Y.-q. Song,
    Impact of Cr(VI) on P removal performance in enhanced
    biological phosphorus removal (EBPR) system based on
    the anaerobic and aerobic metabolism, Bioresour. Technol.,
    121 (2014) 379–385. 
-  S.-b. Xia, J.-x. Liu, An innovative integrated oxidation ditch
    with vertical circle (IODVC) for wastewater treatment, Process
    Biochem., 39 (2004) 1111–1117. 
-  R. Liu, X. Hao, Q. Chen, J. Li, Research advances of Tetrasphaera
    in enhanced biological phosphorus removal: a review, Water
    Res., 166 (2019) 115–122. 
-  F.J. Rubio-Rincón, D.G. Weissbrodt, C.M. Lopez-Vazquez, L. Welles, B. Abbas, M. Albertsen, P.H. Nielsen,
    
 M.C.M. van Loosdrecht, D. Brdjanovic, “Candidatus
    accumulibacter delftensis”: a clade IC novel polyphosphateaccumulating
    organism without denitrifying activity on nitrate,
    Water Res., 161 (2019) 136–151.
-  W. Song, M.J. Zheng, H. Li, W. Zheng, F. Guo, Profiling
    population-level diversity and dynamics of Accumulibacter via
    high throughput sequencing of ppk1, Microbiol. Biotechnol.,
    103 (2019) 9711–9722. 
-  Y.-M. Guo, Y.-G. Liu, G.-M. Zeng, X.-J. Hu, W.-H. Xu, Y.-Q. Liu,
    S.-M. Liu, H.-S. Sun, J. Ye, H.-J. Huang, An integrated treatment
    of domestic wastewater using sequencing batch biofilm reactor
    combined with vertical flow constructed wetland and its
    artificial neural network simulation study, Ecol. Eng., 64 (2014)
    18–26. 
-  G.W. Fuhs, M. Chen, Microbiological basis of phosphate
    removal in the activated sludge process for the treatment of
    wastewater, Microb. Ecol., 2 (1975) 119–138. 
-  L.H. Lötter, The role of bacterial phosphate metabolism in
    enhanced phosphorus removal from the activated sludge
    process, Water Sci. Technol., 17 (1985) 127–138. 
-  Y. Wang, Y. Peng, T. Stephenson, Effect of influent nutrient
    ratios and hydraulic retention time (HRT) on simultaneous
    phosphorus and nitrogen removal in a two-sludge sequencing
    batch reactor process, Bioresour. Technol., 100 (2009) 3506–3512. 
-  W. Zeng, L. Zhang, P. Fan, J. Guo, Y. Peng, Community structures
    and population dynamics of “Candidatus Accumulibacter” in
    activated sludges of wastewater treatment plants using ppk1 as
    phylogenetic marker, Environ. Sci., 67 (2018) 237–248. 
-  X. Huang, W. Dong, H. Wang, S. Jiang, Biological nutrient
    removal and molecular biological characteristics in an
    anaerobic-multistage anaerobic/oxic (A-MAO) process to treat
    municipal wastewater, Bioresour. Technol., 241 (2017) 969–978. 
-  A. Oehmen, R. Keller-Lehmann, R.J. Zeng, Z.G. Yuan, J. Keller,
    Optimisation of poly-beta-hydroxyalkanoate analysis using gas
    chromatography for enhanced biological phosphorus removal
    systems, Chromatography, 1070 (2005) 131–136. 
-  P.L. Bond, R. Erhart, M. Wagner, J. Keller, L.L. Blackall,
    Identification of some of the major groups of bacteria in efficient
    and nonefficient biological phosphorus removal activated
    sludge systems, Environ. Microbiol., 65 (1999) 4077–4084. 
-  G. Sin, K. Niville, G. Bachis, T. Jiang, I. Nopens, S. van Hulle,
    P.A. Vanrolleghem, Nitrite effect on the phosphorus uptake
    activity of phosphate accumulating organisms (PAOs) in pilotscale
    SBR and MBR reactors, Water SA, 34 (2008) 17–38. 
-  Y. Yoshida, Y. Kim, T. Saito, K. Tanaka, Development of the
    modified activated sludge model describing nitrite inhibition
    of aerobic phosphate uptake, Water Sci. Technol., 59 (2009)
    621–637. 
-  H. Li, H. Liu, Q. Zeng, M. Xu, Y. Li, W. Wang, Y. Zhong,
    Isolation and appraisal of a non-fermentative bacterium, Delftia
    tsuruhatensis, as denitrifying phosphate-accumulating organism
    and optimal growth conditions, Water Process Eng., 36 (2020)
    1012–1034. 
-  H. Zou, Y. Wang, Phosphorus removal and recovery from
    domestic wastewater in a novel process of enhanced biological
    phosphorus removal coupled with crystallization, Bioresour.
    Technol., 211 (2021) 87–92. 
-  Z. Lin, Y. Wang, W. Huang, J. Wang, L. Chen, J. Zhou, Q. He,
    Single-stage denitrifying phosphorus removal biofilter utilizing
    intracellular carbon source for advanced nutrient removal and
    phosphorus recovery, Bioresour. Technol., 277 (2019) 27–36. 
-  C. Zhang, Y. Qian, L. Yuan, S. He, Y. Wang, L. Wang, Nutrients
    removal performance of a denitrifying phosphorus removal
    process in alternate anaerobic/anoxic-aerobic double membrane
    bioreactors 
 (A2N-DMBR), Water Sci. Technol., 78 (2018)
    1741–1752.
-  H. Liu, Q. Wang, Y. Sun, K. Zhou, W. Liu, Q. Lu, C. Ming,
    X. Feng, J. Du, X. Jia, J. Li, solation of a non-fermentative
    bacterium, Pseudomonas aeruginosa, using intracellular carbon
    for denitrification and
 phosphorus-accumulation and relevant
    metabolic mechanisms, Bioresour. Technol., 211 (2016) 6–15.
-  Y. Barak, J. van Rijn, Relationship between nitrite reduction
    and active phosphate uptake in the phosphate-accumulating
    denitrifier Pseudomonas sp. strain JR 12, Environ. Microbiol.,
    66 (2000) 5236–5240. 
-  H. Wang, Q. He, D. Chen, L. Wei, Z. Zou, J. Zhou, K. Yang,
    H. Zhang, Microbial community in a hydrogenotrophic
    denitrification reactor based on pyrosequencing, Microbiol.
    Biotechnol., 99 (2015) 10829–10837. 
-  H. Yu, M. Li, Denitrifying and phosphorus accumulating
    mechanisms of denitrifying phosphorus accumulating
    organisms (DPAOs) for wastewater treatment—a review, Acta
    Microbiol. Sin., 55 (2015) 264–272. 
-  M. Zhang, J. Gao, Q. Liu, Y. Fan, C. Zhu, Y. Liu, C. He,
    J. Wu, Nitrite accumulation and microbial behavior by
    seeding denitrifying phosphorus removal sludge for partial
	  denitrification (PD): the effect of COD/NO3– ratio, Bioresour.
  Technol., 323 (2020) 1245–1257. 
-  C. Li, J. Zhang, S. Liang, H.H. Ngo, W. Guo, Y. Zhang, Y. Zou,
    Nitrous oxide generation in denitrifying phosphorus removal
    process: main causes and control measures, Environ. Sci. Pollut.
    Res., 20 (2013) 5353–5360. 
-  Y. Sun, Y. Peng, J. Zhang, X. Li, Q. Zhang, L. Zhang, Effect
    of endogenous metabolisms on survival and activities of
    denitrifying phosphorus removal sludge under various
    starvation conditions, Bioresour. Technol., 315 (2020) 123839,
    doi: 10.1016/j.biortech.2020.123839. 
-  Q. Zhao, M. Yu, H. Lu, Y.-H. Zhang, B.K. Biswal, G.-H. Chen,
    D. Wu, Formation and characterization of the 
 micro-size
    granular sludge in denitrifying sulfur-conversion associated
    enhanced biological phosphorus removal (DS-EBPR)
    process, Bioresour. Technol., 291 (2019) 121871, doi: 10.1016/j.
    biortech.2019.121871.
-  T. Zhang, M.-F. Shao, L. Ye, 454 pyrosequencing reveals bacterial
    diversity of activated sludge from 14 sewage treatment plants,
    ISME J., 62 (2012) 1137–1147. 
-  Q. Ma, Y. Qu, W. Shen, Z. Zhang, J. Wang, Z. Liu, D. Li,
    H. Li, J. Zhou, Bacterial community compositions of coking
    wastewater treatment plants in steel industry revealed by
    Illumina high-throughput sequencing, Bioresour. Technol.,
    179 (2015) 436–443. 
-  R. Du, S. Cao, B. Li, M. Niu, S. Wang, Y. Peng, Performance and
    microbial community analysis of a novel DEAMOX based on
    partial-denitrification and anammox treating ammonia and
    nitrate wastewaters, Water Res., 108 (2017) 46–56. 
-  C. Wan, Q. Zhang, D.-J. Lee, Y. Wang, J. Li, Long-term storage
    of aerobic granules in liquid media: viable but non-culturable
    status, Bioresour. Technol., 166 (2014) 464–470. 
-  Z. Jin, F.-Y. Ji, X. Xu, X.-Y. Xu, Q.-K. Chen, Q. Li, Microbial
    and metabolic characterization of a denitrifying phosphorusuptake/
    side stream phosphorus removal system for treating
    domestic sewage, Biodegradation, 25 (2014) 777–786. 
-  M.F. Carosia, D.Y. Okada, I.K. Sakamoto, E.L. Silva,
    M.B. Amâncio Varesche, Microbial characterization and
    degradation of linear alkylbenzene sulfonate in an anaerobic
    reactor treating wastewater containing soap powder,
    Bioresour. Technol., 167 (2014) 316–323. 
-  Q. He, J. Zhou, H. Wang, J. Zhang, L. Wei, Microbial population
    dynamics during sludge granulation in an A/O/A sequencing
    batch reactor, Bioresour. Technol., 214 (2016) 1–8. 
-  R. Liao, Y. Li, J. Du, A. Li, H. Song, Z. Shen, Y. Li, Analysis of
    high-nitrate, high-salinity wastewater in an expanded granular
    sludge bed reactor and microbial community, Desal. Water
    Treat., 57 (2016) 4357–4364. 
-  T. Shigematsu, K. Yumihara, Y. Ueda, M. Numaguchi,
    S. Morimura, K. Kida, Delftia tsuruhatensis sp. nov., 
 a
    terephthalate-assimilating bacterium isolated from activated
    sludge, Syst. Evol. Microbiol., 53 (2019) 1479–1483.